Comments: Placed in a separate subtribe, Macacina, from other members of the Papionini, by McKenna and Bell (1997). Species groups are: (1) M. sylvanus group (monotypic), (2) M. nemestrina group (silenus, leonina, nemestrina, pagensis, siberu, maura, ochreata, tonkeana, hecki, nigrescens, nigra), (3) M. fascicularis group (fascicularis, arctoides), (4) M. mulatta group (mulatta, cyclopis, fuscata), (5) M. sinica group (sinica, radiata, assamensis, thibetana)